Scientists have found that it was not the dinosaurs who ruled Earth 13 million years ago in Peru, it was the crocodiles.

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There were at least seven different species of the reptile that hunted in the lakes, swamps and rivers of the massive wetland region that pre-dated the Amazon basin, the Mirror reported. Scientists uncovered the crocodile fossils from a rock layer known as the Pebas Formation in north-eastern Peru in a series of excavations conducted since 2002, of which three of the species are entirely new to science.

Gnatusuchus pebasensis, a short-faced caiman with "globular teeth" is the most unusual, which is believed to have used its snout as a shovel to dig for clams and other mollusks. These species vanished when the mega-wetlands transformed into the modern Amazon and mollusk numbers and diversity declined.

Researcher Rodolfo Salas-Gismondi, from the University of Montpellier in France, said that they "uncovered this special moment in time when the ancient mega-wetland ecosystem reached its peak in size and complexity, just before its demise and the start of the modern Amazon River system".

It's a milestone for understanding proto-Amazonian wetland feeding dynamics, he added. The team also found the first clear fossil representative of modern smooth-fronted caimans which was adapted to catching a variety of prey including fish. Today, six caiman species live in the whole Amazon basin but only three ever co-exist in the same area and they rarely share the same habitat.

The findings are described in the journal Proceedings of the Royal Society B.